SBMNH Entomology - The California Beetle Database

Family	Genus	Species	Author	Intro?	No. Records
Scarabaeidae	Acoma	arizonica	Brown	N	4
Scarabaeidae	Acoma	glabrata	Cazier	N	4
Scarabaeidae	Aegialia	blanchardi	Horn	N	4
Scarabaeidae	Aegialia	browni	Saylor	N	9
Scarabaeidae	Aegialia	concinna	Gordon & Cartwright	N	7
Scarabaeidae	Aegialia	conferta	Horn	N	3
Scarabaeidae	Aegialia	convexa	Fall	N	44
Scarabaeidae	Aegialia	crassa	LeConte	N	8
Scarabaeidae	Aegialia	cylindrica	(Eschscholtz)	N	1
Scarabaeidae	Aegialia	kelsoi	Gordon & Cartwright	N	1
Scarabaeidae	Aegialia	lacustris	LeConte	N	1
Scarabaeidae	Aegialia	latispina	LeConte	N	35
Scarabaeidae	Aegialia	nigrella	Brown	N	2
Scarabaeidae	Aegialia	opaca	Brown	N	1
Scarabaeidae	Aegialia	punctata	Brown	N	4
Scarabaeidae	Aegialia	spinosa	Gordon & Cartwright	N	1
Scarabaeidae	Annegialia	ataeniformis	Howden	N	1
Scarabaeidae	Anomala	carlsoni	Hardy	N	16
Scarabaeidae	Anomala	flavilla	Bates	N	13
Scarabaeidae	Anomala	hardyorum	Potts	N	2
Scarabaeidae	Anomala	imperialiae	Potts	N	2
Scarabaeidae	Aphodius	aculeatus	Robinson	N	1
Scarabaeidae	Aphodius	acutissimus	Gordon	N	3
Scarabaeidae	Aphodius	aleutus	Eschscholtz	N	1
Scarabaeidae	Aphodius	alternatus	Horn	N	45
Scarabaeidae	Aphodius	cadaverinus	(Mannerheim)	N	1
Scarabaeidae	Aphodius	carpinterius	Gordon & Skelley	N	3
Scarabaeidae	Aphodius	coloradensis	Horn	N	1
Scarabaeidae	Aphodius	congregatus	Mannerheim	N	1
Scarabaeidae	Aphodius	consociatus	Horn	N	3
Scarabaeidae	Aphodius	conspersus	Horn	N	2
Scarabaeidae	Aphodius	coquilletti	Linell	N	2
Scarabaeidae	Aphodius	crenicollis	Fall	N	1
Scarabaeidae	Aphodius	cribratulus	Schmidt	N	1
Scarabaeidae	Aphodius	davisi	Fall	N	64
Scarabaeidae	Aphodius	denticulatus	Haldeman	N	1
Scarabaeidae	Aphodius	distinctus	(Muller)	Y	1
Scarabaeidae	Aphodius	essigi	Saylor	N	1
Scarabaeidae	Aphodius	fimetarius	(Linnaeus)	Y	78
Scarabaeidae	Aphodius	fossor	(Linnaeus)	Y	1
Scarabaeidae	Aphodius	fucosus	Schmidt	N	10
Scarabaeidae	Aphodius	gentilis	Horn	N	1
Scarabaeidae	Aphodius	granarius	(Linnaeus)	Y	65
Scarabaeidae	Aphodius	gravis	Fall	N	1
Scarabaeidae	Aphodius	haemorrhoidalis	(Linnaeus)	Y	9
Scarabaeidae	Aphodius	hamatus	Say	N	4
Scarabaeidae	Aphodius	inyoensis	Van Dyke	N	1
Scarabaeidae	Aphodius	linsleyi	Van Dyke	N	1
Scarabaeidae	Aphodius	lividus	(Olivier)	Y	68
Scarabaeidae	Aphodius	luxatus	Horn	N	22
Scarabaeidae	Aphodius	martini	Van Dyke	N	1
Scarabaeidae	Aphodius	militaris	LeConte	N	106
Scarabaeidae	Aphodius	neotomae	Fall	N	2
Scarabaeidae	Aphodius	nevadensis	Horn	N	1
Scarabaeidae	Aphodius	ochreipennis	Horn	N	1
Scarabaeidae	Aphodius	opacus	LeConte	N	58
Scarabaeidae	Aphodius	oviformis	Robinson	N	1
Scarabaeidae	Aphodius	ovipennis	Horn	N	41
Scarabaeidae	Aphodius	pardalis	LeConte	N	4
Scarabaeidae	Aphodius	pectoralis	LeConte	N	1
Scarabaeidae	Aphodius	pedellus	(DeGeer)	N	1
Scarabaeidae	Aphodius	pseudofucosus	Gordon & Skelley	N	12
Scarabaeidae	Aphodius	ruficlarus	Fall	N	8
Scarabaeidae	Aphodius	rugatus	Schmidt	N	21
Scarabaeidae	Aphodius	rugoclypeus	Hinton	N	1
Scarabaeidae	Aphodius	slevini	Van Dyke	N	1
Scarabaeidae	Aphodius	sparsus	LeConte	N	54
Scarabaeidae	Aphodius	spermophili	Gordon	N	2
Scarabaeidae	Aphodius	subaeneus	LeConte	N	25
Scarabaeidae	Aphodius	tuberosus	Barrett	N	7
Scarabaeidae	Aphodius	ungulatus	Fall	N	3
Scarabaeidae	Aphodius	vandykei	Barret	N	90
Scarabaeidae	Aphodius	vittatus	Say	N	10
Scarabaeidae	Ataenius	californicus	Horn	N	30
Scarabaeidae	Ataenius	cognatus	(LeConte)	N	10
Scarabaeidae	Ataenius	desertus	Horn	N	28
Scarabaeidae	Ataenius	duncani	Cartwright	N	6
Scarabaeidae	Ataenius	hesperius	Cartwright	N	22
Scarabaeidae	Ataenius	lobatus	Horn	N	16
Scarabaeidae	Ataenius	nocturnus	(Nomura)	N	25
Scarabaeidae	Ataenius	platensis	(Blanchard)	N	7
Scarabaeidae	Ataenius	puncticollis	(LeConte)	N	6
Scarabaeidae	Ataenius	punctifrons	Cartwright	N	4
Scarabaeidae	Ataenius	sculptor	Harold	N	1
Scarabaeidae	Ataenius	spretulus	(Haldeman)	N	1
Scarabaeidae	Ataenius	stephani		N	5
Scarabaeidae	Ataenius	vandykei	Cartwright	N	1
Scarabaeidae	Bolbocerastes	regalis		N	4
Scarabaeidae	Canthon	simplex	LeConte	N	34
Scarabaeidae	Chnaunanthus	chapini	Saylor	N	2
Scarabaeidae	Chnaunanthus	discolor	Burmeister	N	2
Scarabaeidae	Chnaunanthus	flavipennis	Horn	N	3
Scarabaeidae	Coenonycha	acuta	Cazier	N	7
Scarabaeidae	Coenonycha	ampla	Cazier	N	14
Scarabaeidae	Coenonycha	barri	Cazier	N	3
Scarabaeidae	Coenonycha	bowlesi	Cazier	N	3
Scarabaeidae	Coenonycha	clementina	Casey	N	14
Scarabaeidae	Coenonycha	clypeata	McClay	N	10
Scarabaeidae	Coenonycha	crispata	McClay	N	5
Scarabaeidae	Coenonycha	dimorpha	Evans	N	15
Scarabaeidae	Coenonycha	fuga	Cazier	N	4
Scarabaeidae	Coenonycha	fulva	McClay	N	22
Scarabaeidae	Coenonycha	fusca	McClay	N	6
Scarabaeidae	Coenonycha	hageni	Cazier	N	4
Scarabaeidae	Coenonycha	lurida	Cazier	N	5
Scarabaeidae	Coenonycha	mediata	Cazier	N	2
Scarabaeidae	Coenonycha	ochreata	Evans	N	8
Scarabaeidae	Coenonycha	ovatis	McClay	N	3
Scarabaeidae	Coenonycha	pallida	Cazier	N	10
Scarabaeidae	Coenonycha	parvula	Fall	N	5
Scarabaeidae	Coenonycha	pascuensis	Potts	N	2
Scarabaeidae	Coenonycha	purshiae	Cazier	N	3
Scarabaeidae	Coenonycha	rotundata	(LeConte)	N	9
Scarabaeidae	Coenonycha	santacruzae	Evans	N	4
Scarabaeidae	Coenonycha	saylori	Cazier	N	2
Scarabaeidae	Coenonycha	scotti	McClay	N	9
Scarabaeidae	Coenonycha	testacea	Cazier	N	43
Scarabaeidae	Coenonycha	tingi	Cazier	N	12
Scarabaeidae	Cotalpa	flavida	Horn	N	1
Scarabaeidae	Cotinis	mutabilis	(Gory & Percheron)	N	8
Scarabaeidae	Cremastocheilus	angularis	LeConte	N	8
Scarabaeidae	Cremastocheilus	armatus	Walker	N	22
Scarabaeidae	Cremastocheilus	armatus armatus	Walker	N	1
Scarabaeidae	Cremastocheilus	armatus montanus		N	16
Scarabaeidae	Cremastocheilus	beameri	Cazier	N	1
Scarabaeidae	Cremastocheilus	crinitus	LeConte	N	1
Scarabaeidae	Cremastocheilus	depressus	Horn	N	2
Scarabaeidae	Cremastocheilus	opaculus	Horn	N	1
Scarabaeidae	Cremastocheilus	planatus	LeConte	N	2
Scarabaeidae	Cremastocheilus	puncticollis	Cazier	N	1
Scarabaeidae	Cremastocheilus	quadratus	Fall	N	3
Scarabaeidae	Cremastocheilus	schaumii	LeConte	N	11
Scarabaeidae	Cremastocheilus	schaumii	LeConte	N	3
Scarabaeidae	Cremastocheilus	schaumii schaumii	LeConte	N	2
Scarabaeidae	Cremastocheilus	schaumii westwoodi	Horn	N	1
Scarabaeidae	Cremastocheilus	westwoodi	Horn	N	8
Scarabaeidae	Cremastocheilus	westwoodi tridens	Casey	N	1
Scarabaeidae	Cremastocheilus	westwoodi westwoodi	Horn	N	2
Scarabaeidae	Cremastocheilus	wheeleri	LeConte	N	2
Scarabaeidae	Cyclocephala	hirta	LeConte	N	22
Scarabaeidae	Cyclocephala	longula	LeConte	N	63
Scarabaeidae	Cyclocephala	melanocephala	(Fabricius)	N	17
Scarabaeidae	Cyclocephala	pasadenae	(Casey)	N	5
Scarabaeidae	Cyclocephala	pilosicollis	(Saylor)	N	1
Scarabaeidae	Cyclocephala	wandae	Hardy	N	3
Scarabaeidae	Dichelonyx	backii		N	6
Scarabaeidae	Dichelonyx	clypeata	(Horn)	N	1
Scarabaeidae	Dichelonyx	decolorata	(Fall)	N	1
Scarabaeidae	Dichelonyx	fulgida	LeConte	N	43
Scarabaeidae	Dichelonyx	lateralis	(Fall)	N	1
Scarabaeidae	Dichelonyx	muscula	(Fall)	N	14
Scarabaeidae	Dichelonyx	nana	(Fall)	N	4
Scarabaeidae	Dichelonyx	pallens	LeConte	N	1
Scarabaeidae	Dichelonyx	pusilla	LeConte	N	84
Scarabaeidae	Dichelonyx	robusta	(Fall)	N	4
Scarabaeidae	Dichelonyx	truncata	LeConte	N	46
Scarabaeidae	Dichelonyx	vaga	(Fall)	N	9
Scarabaeidae	Dichelonyx	valida	LeConte	N	1
Scarabaeidae	Dichelonyx	vandykei	Saylor	N	1
Scarabaeidae	Dichelonyx	vicina	(Fall)	N	61
Scarabaeidae	Digitonthophagus	gazella	(Fabricius)	Y	1
Scarabaeidae	Dinacoma	caseyi	Blaisdell	N	8
Scarabaeidae	Dinacoma	marginata	(Casey)	N	31
Scarabaeidae	Diplotaxis	anxius	LeConte	N	6
Scarabaeidae	Diplotaxis	brevicollis	LeConte	N	1
Scarabaeidae	Diplotaxis	conformis	Fall	N	1
Scarabaeidae	Diplotaxis	corbula	Vaurie	N	22
Scarabaeidae	Diplotaxis	corvina	LeConte	N	4
Scarabaeidae	Diplotaxis	dahli	Cazier	N	1
Scarabaeidae	Diplotaxis	deserta	Fall	N	3
Scarabaeidae	Diplotaxis	fimbriata	Fall	N	27
Scarabaeidae	Diplotaxis	fossipalpa	Fall	N	16
Scarabaeidae	Diplotaxis	haydeni	LeConte	N	1
Scarabaeidae	Diplotaxis	incuria	Fall	N	1
Scarabaeidae	Diplotaxis	insignis	LeConte	N	18
Scarabaeidae	Diplotaxis	knausii	Schaeffer	N	40
Scarabaeidae	Diplotaxis	magna	Bates	N	1
Scarabaeidae	Diplotaxis	missionaria	Vaurie	N	3
Scarabaeidae	Diplotaxis	moerens	LeConte	N	50
Scarabaeidae	Diplotaxis	obscura	LeConte	N	9
Scarabaeidae	Diplotaxis	pacata	LeConte	N	18
Scarabaeidae	Diplotaxis	rufiola	Fall	N	5
Scarabaeidae	Diplotaxis	sierrae	Fall	N	85
Scarabaeidae	Diplotaxis	subangulata	LeConte	N	58
Scarabaeidae	Diplotaxis	tenuis	LeConte	N	2
Scarabaeidae	Euoniticellus	intermedius	(Reiche)	Y	1
Scarabaeidae	Euphoria	fascifera trapezium	Casey	N	1
Scarabaeidae	Genuchinus	ineptus	(Horn)	N	1
Scarabaeidae	Gymnopyge	coquilletti	Linell	N	1
Scarabaeidae	Gymnopyge	hirsuta	Cazier	N	1
Scarabaeidae	Gymnopyge	hopliaeformis	Linell	N	1
Scarabaeidae	Hemiphileurus	illatus	(LeConte)	N	7
Scarabaeidae	Hoplia	callipyge	LeConte	N	492
Scarabaeidae	Hoplia	dispar	LeConte	N	211
Scarabaeidae	Hoplia	hirta	LeConte	N	8
Scarabaeidae	Hoplia	sackenii	LeConte	N	100
Scarabaeidae	Leptohoplia	testaceipennis	Saylor	N	4
Scarabaeidae	Liatongus	californicus	(Horn)	N	2
Scarabaeidae	Megasoma	sleeperi	Hardy	N	6
Scarabaeidae	Neopsammodius	blandus	(Fall)	N	1
Scarabaeidae	Neopsammodius	quinqueplicatus	(Horn)	N	1
Scarabaeidae	Oncerus	floralis	LeConte	N	21
Scarabaeidae	Onitis	alexis	Klug	Y	5
Scarabaeidae	Onthophagus	cartwrighti	Howden	N	1
Scarabaeidae	Onthophagus	taurus	(Schreber)	Y	21
Scarabaeidae	Onthophagus	velutinus	Horn	N	3
Scarabaeidae	Oxygrylius	peninsularis	Casey	N	1
Scarabaeidae	Oxygrylius	ruginasus	(LeConte)	N	30
Scarabaeidae	Paracotalpa	deserta	Saylor	N	14
Scarabaeidae	Paracotalpa	granicollis	(Haldeman)	N	2
Scarabaeidae	Paracotalpa	puncticollis	(LeConte)	N	11
Scarabaeidae	Paracotalpa	ursina	(Horn)	N	37
Scarabaeidae	Parathyce	bidentata	(Fall)	N	3
Scarabaeidae	Parathyce	carpenteri	(LeConte)	N	47
Scarabaeidae	Parathyce	fieldi	(Fall)	N	37
Scarabaeidae	Parathyce	harfordi	(Casey)	N	25
Scarabaeidae	Parathyce	palpalis	(Horn)	N	153
Scarabaeidae	Parathyce	riversi	(Casey)	N	21
Scarabaeidae	Phobetus	ciliatus	Barrett	N	12
Scarabaeidae	Phobetus	comatus	LeConte	N	58
Scarabaeidae	Phobetus	humeralis	Cazier	N	58
Scarabaeidae	Phobetus	leachi	Barrett	N	1
Scarabaeidae	Phobetus	mojavus	Barrett	N	17
Scarabaeidae	Phobetus	palpalis	Saylor	N	8
Scarabaeidae	Phobetus	panamintensis	Hardy	N	3
Scarabaeidae	Phobetus	robinsoni	Saylor	N	5
Scarabaeidae	Phobetus	saylori	Cazier	N	14
Scarabaeidae	Phobetus	testaceus	LeConte	N	18
Scarabaeidae	Phyllophaga	anxia	(LeConte	N	1
Scarabaeidae	Phyllophaga	errans	(LeConte)	N	2
Scarabaeidae	Phyllophaga	falsa	(LeConte)	N	1
Scarabaeidae	Phyllophaga	fervida	(Fabricius)	N	1
Scarabaeidae	Phyllophaga	mucorea	(LeConte)	N	6
Scarabaeidae	Phyllophaga	reevesi	Saylor	N	1
Scarabaeidae	Phyllophaga	sequoiana	Saylor	N	1
Scarabaeidae	Phyllophaga	stohleri	Saylor	N	2
Scarabaeidae	Phyllophaga	timida	(Horn)	N	1
Scarabaeidae	Phyllophaga	xerophila	Saylor	N	1
Scarabaeidae	Platyomus	grundeli	(Van Dyke)	N	1
Scarabaeidae	Platyomus	micros	(Bates)	N	1
Scarabaeidae	Platyomus	tibialis	(Fabricius)	N	11
Scarabaeidae	Plectrodes	pubescens	Horn	N	28
Scarabaeidae	Pleurophorus	caesus	(Creutzer)	N	3
Scarabaeidae	Polyphylla	anteronivea	Hardy & Andrews	N	1
Scarabaeidae	Polyphylla	arguta	Casey	N	1
Scarabaeidae	Polyphylla	barbata	Cazier	N	3
Scarabaeidae	Polyphylla	cavifrons	LeConte	N	11
Scarabaeidae	Polyphylla	crinita	LeConte	N	11
Scarabaeidae	Polyphylla	decemlineata	(Say)	N	41
Scarabaeidae	Polyphylla	diffracta	Casey	N	1
Scarabaeidae	Polyphylla	erratica	Hardy & Andrews	N	3
Scarabaeidae	Polyphylla	modulata	Casey	N	1
Scarabaeidae	Polyphylla	nigra	Casey	N	11
Scarabaeidae	Polyphylla	nubila	Van Dyke	N	1
Scarabaeidae	Polyphylla	rugosipennis	Casey	N	1
Scarabaeidae	Polyphylla	sobrina	Casey	N	5
Scarabaeidae	Polyphylla	stellata	Young	N	1
Scarabaeidae	Pseudocotalpa	andrewsi	Hardy	N	7
Scarabaeidae	Pseudocotalpa	giulianii	Hardy	N	3
Scarabaeidae	Rhyssemus	californicus	Horn	N	34
Scarabaeidae	Serica	abdita	Dawson	N	1
Scarabaeidae	Serica	acicula	Dawson	N	5
Scarabaeidae	Serica	acontia	Dawson	N	3
Scarabaeidae	Serica	adversa	Dawson	N	3
Scarabaeidae	Serica	alleni	Saylor	N	21
Scarabaeidae	Serica	alternata	LeConte	N	80
Scarabaeidae	Serica	alternata alternata	LeConte	N	11
Scarabaeidae	Serica	alternata exolita	Dawson	N	7
Scarabaeidae	Serica	alternata patruela	Dawson	N	1
Scarabaeidae	Serica	anthracina	LeConte	N	15
Scarabaeidae	Serica	calignosa	Dawson	N	1
Scarabaeidae	Serica	catalina	Dawson	N	3
Scarabaeidae	Serica	chaetosoma	Dawson	N	5
Scarabaeidae	Serica	chicoensis	Saylor	N	1
Scarabaeidae	Serica	coalinga	Dawson	N	1
Scarabaeidae	Serica	cruzi	Saylor	N	4
Scarabaeidae	Serica	curvata	LeConte	N	13
Scarabaeidae	Serica	cuyamaca	Saylor	N	2
Scarabaeidae	Serica	deserticola	Dawson	N	1
Scarabaeidae	Serica	diablo	Dawson	N	25
Scarabaeidae	Serica	egregia	Dawson	N	9
Scarabaeidae	Serica	elmontea	Saylor	N	17
Scarabaeidae	Serica	elongatula	Horn	N	1
Scarabaeidae	Serica	falcata	Dawson	N	1
Scarabaeidae	Serica	falli	Dawson	N	1
Scarabaeidae	Serica	fimbriata	LeConte	N	30
Scarabaeidae	Serica	heteracantha	Dawson	N	6
Scarabaeidae	Serica	laguna	Saylor	N	1
Scarabaeidae	Serica	ligulata	Dawson	N	46
Scarabaeidae	Serica	ligulata ligulata	Dawson	N	9
Scarabaeidae	Serica	ligulata praetermissa	Dawson	N	28
Scarabaeidae	Serica	mckenziei	Saylor	N	8
Scarabaeidae	Serica	mixta	LeConte	N	6
Scarabaeidae	Serica	oliver	Saylor	N	2
Scarabaeidae	Serica	pavonia	Dawson	N	1
Scarabaeidae	Serica	perigonia	Dawson	N	20
Scarabaeidae	Serica	porcula	Casey	N	4
Scarabaeidae	Serica	prava	Dawson	N	8
Scarabaeidae	Serica	pruinipennis	Saylor	N	2
Scarabaeidae	Serica	pullata	Dawson	N	1
Scarabaeidae	Serica	repanda	Dawson	N	1
Scarabaeidae	Serica	rossi	Saylor	N	1
Scarabaeidae	Serica	sandiegensis	Saylor	N	2
Scarabaeidae	Serica	satrapa	Dawson	N	5
Scarabaeidae	Serica	scaphia	Dawson	N	1
Scarabaeidae	Serica	sculptilis	Dawson	N	31
Scarabaeidae	Serica	searli	Saylor	N	1
Scarabaeidae	Serica	senta	Dawson	N	1
Scarabaeidae	Serica	serotina	LeConte	N	7
Scarabaeidae	Serica	solita	Dawson	N	1
Scarabaeidae	Serica	stygia	Dawson	N	10
Scarabaeidae	Serica	subnisa	Dawson	N	1
Scarabaeidae	Serica	ventura dorsalis	Dawson	N	1
Scarabaeidae	Serica	ventura personata	Dawson	N	1
Scarabaeidae	Serica	ventura ventura	Dawson	N	1
Scarabaeidae	Serica	watson	Saylor	N	1
Scarabaeidae	Tesarius	caelatus	(LeConte)	N	5
Scarabaeidae	Tesarius	doyeni	(Cartwright)	N	7
Scarabaeidae	Tesarius	mcclayi	(Cartwright)	N	1
Scarabaeidae	Tesarius	oregonensis	(Cartwright)	N	1
Scarabaeidae	Thyce	deserta	Hardy	N	12
Scarabaeidae	Tomarus	gibbosus obsoletus	(LeConte)	N	44
Scarabaeidae	Trichiorhyssemus	riparius	(Horn)	N	4
Scarabaeidae	Valgus	californicus	Horn	N	12
Scarabaeidae	Xeropsamobeus	desertus	(Van Dyke)	N	12



	Collections and Research Online Databases - California Beetle Project